The 335-hp V8 engine can propel the GL 450 from a standing start to 60 miles per hour in a car-like 7.4 seconds. This powerplant, with four valves per cylinder, is part of a brand new family of V8 engines from Mercedes-Benz.

A standard, seven-speed automatic transmission helps keep the engine operating in the sweet part of a power band that provides maximum torque from 2700 rpm all the way up to 5000. Mercedes notes that the GL is the only full-size SUV offering such a seven-speed gearbox.

The 4MATIC four-wheel-drive system features front, center and rear differentials. The sure-footed 4MATIC system is designed to maintain mobility even when only wheel has traction. In normal driving, the system distributes power equally to front and rear wheels. The system includes DSR, a sort of cruise control for regulating downhill speed, and Hill-Start Assist, which keeps the GL from rolling backward when launched after being stopped on uphill slope. 4MATIC also adjusts anti-lock brake controls to provide quicker stops on slippery and unpaved surfaces when off-roading.

Airmatic is an air suspension system that uses air bladders instead of coil springs to adjust ride height by as much as three inches as well as ride firmness and body control parameters, so the driver can select from comfort and more aggressively sporty settings. The system automatically lowers the vehicle to enhance handling and aerodynamic efficiency at speeds of 77 mph or higher.

Those serious about venturing far from pavement can opt for a special off-road package that includes special wheels and tires, a two-speed transfer case, locking center and rear differentials, adaptive dampening and with ground clearance height-adjustment options of as much as 12.4 inches, compared to the standard 7.9 inches of clearance.

We did our driving on freeways in and out of San Francisco's airport and on roads through Napa Valley wine country. Steering feedback was consistent, brakes (14-inches in diameter on the front wheels and 13 inches on the rear) were responsive and consistently predictable and the GL 450 offered nice balanced dynamic capabilities when hustled through the curves on the narrow, hilly roads that wind through the vine-covered hillsides.